Normally, that would be a big, fat "no!". But given the ability to push/tow start these old automatic Benzes I have to ask.

It wasn't referenced in the owner's manual but I discovered by accident that mine does, indeed, start in gear.

$150+ for a new one if mine's busted or has been bypassed.

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1974 240D "Boldie" 170K.- New timing chain/freshly rebuilt IP/replaced valve seals/injectors/upgraded stereo/new Bilsteins with Yokohamas/fresh paint and rocker panels plus lots of welds.